chapter vii

.) is devoted to the understanding, its origin and that of ideas. The following additions relative to chapters vii. and viii. of the first part of this work are from vol. ii., pp. 451-466.

In the last of June, 1809, the menagerie of the Museum of Natural History having received a Phoca (_Phoca vitulina_), Lamarck, as he says, had the opportunity of observing its movements and habits. After describing its habits in swimming and moving on land and observing its relation to the clawed mammals, he says his main object is to remark that the seals do not have the hind legs arranged in the same direction as the axis of their body, because these animals are constrained to habitually use them to form a caudal fin, closing and widening, by spreading their digits, the paddle (_palette_) which results from their union.

"The morses, on the contrary, which are accustomed to feed on grass near the shore, never use their hind feet as a caudal fin; but their feet are united together with the tail, and cannot separate. Thus in animals of similar origin we see a new proof of the effect of habits on the form and structure of organs."

He then turns to the flying mammals, such as the flying squirrel (_Sciurus volans_, _ærobates_, _petaurista_, _sagitta_, and _volucella_), and then explains the origin of their adaptation for flying leaps.

"These animals, more modern than the seals, having the habit of extending their limbs while leaping to form a sort of _parachute_, can _only_ make a very prolonged leap when they glide down from a tree or spring only a short distance from one tree to another. Now, by frequent repetitions of such leaps, in the individuals of these races the skin of their sides is expanded on each side into a loose membrane, which connects the hind and fore legs, and which, enclosing a volume of air, prevents their sudden falling. These animals are, moreover, without membranes between the fingers and toes.

"The Galeopithecus (_Lemur volans_), undoubtedly a more ancient form but with the same habits as the flying squirrel (_Pteromys Geoff._), has the skin of the _flancs_ more ample, still more developed, connecting not only the hinder with the fore legs, but in addition the fingers and the tail with the hind feet. Moreover, they leap much farther than the flying squirrels, and even make a sort of flight.[189]

"Finally, the different bats are probably mammals still older than the Galeopithecus, in the habit of extending their membrane and even their fingers to encompass a greater volume of air, so as to sustain their bodies when they fly out into the air.

"By these habits, for so long a period contracted and preserved, the bats have obtained not only lateral membranes, but also an extraordinary elongation of the fingers of their fore feet (with the exception of the thumb), between which are these very ample membranes uniting them; so that these membranes of the hands become continuous with those of the flanks, and with those which connect the tail with the two hind feet, forming in these animals great membranous wings with which they fly perfectly, as everybody knows.

"Such is then the power of habits, which have a singular influence on the conformation of parts, and which give to the animals which have for a long time contracted certain of them, faculties not found in other animals.

"As regards the amphibious animals of which I have often spoken, it gives me pleasure to communicate to my readers the following reflections which have arisen from an examination of all the objects which I have taken into consideration in my studies, and seen more and more to be confirmed.

"I do not doubt but that the mammals have in reality originated from them, and that they are the veritable cradle (_berceau_) of the entire animal kingdom.

"Indeed, we see that the least perfect animals (and they are the most numerous) live only in the water; hence it is probable, as I have said (vol. ii., p. 85), that it is only in the water or in very humid places that nature causes and still forms, under favorable conditions, direct or spontaneous generations which have produced the simplest animalcules and those from which have successively been derived all the other animals.

"We know that the Infusoria, the polyps, and the Radiata only live in the water; that the worms even only live some in the water and others in very damp places.

"Moreover, regarding the worms, which seem to form an initial branch of the animal scale, since it is evident that the Infusoria form another branch, we may suppose that among those of them which are wholly aquatic--namely, which do not live in the bodies of other animals, such as the Gordius and many others still unknown--there are doubtless a great many different aquatic forms; and that among these aquatic worms, those which afterwards habitually expose themselves to the air have probably produced amphibious insects, such as the mosquitoes, the ephemeras, etc., etc., which have successively given origin to all the insects which live solely in the air. But several races of these having changed their habits by the force of circumstances, and having formed habits of a life solitary, retired, or hidden, have given rise to the arachnides, almost all of which also live in the air.

"Finally, those of the arachnides which have frequented the water, which have consequently become progressively habituated to live in it, and which finally cease to expose themselves to the air--this indicates the relations which, connecting the Scolopendræ to Julus, this to the Oniscus, and the last to Asellus, shrimps, etc., have caused the existence of all the Crustacea.

"The other aquatic worms which are never exposed to the air, multiplying and diversifying their races with time, and gradually making progress in the complication of their structure, have caused the formation of the Annelida, Cirripedia, and molluscs, which together form an uninterrupted portion of the animal scale.

"In spite of the considerable hiatus which we observe between the known molluscs and the fishes, the molluscs, whose origin I have just indicated, have, by the intermediation of those yet remaining unknown, given origin to the fishes, as it is evident that the latter have given rise to the reptiles.

"In continuing to consult the probabilities on the origin of different animals, we cannot doubt but that the reptiles, by two distinct branches which circumstances have brought about, have given rise on one side to the formation of birds, and on the other to that of amphibious mammals, which have given in their turn origin to all the other mammals.[190]

"Indeed, the fishes having caused the formation of Batrachia, and these of the Ophidian reptiles, both having only one auricle in the heart, nature has easily come to give a heart with a double auricle to other reptiles which constitute two special branches; finally, she has easily arrived at the end of forming, in the animals which had originated from each of these branches, a heart with two ventricles.

"Thus, among the reptiles whose heart has a double auricle, on the one side, the Chelonians seem to have given origin to the birds; if, independently of several relations which we cannot disregard, I should place the head of a tortoise on the neck of certain birds, I should perceive almost no disparity in the general physiognomy of the factitious animal; and on the other side, the saurians, especially the 'planicaudes,' such as the crocodiles, seem to have given origin to the amphibious mammals.

"If the branch of the Chelonians has given rise to birds, we can yet presume that the palmipede aquatic birds, especially the _brevipennes_, such as the penguins and the _manchots_, have given origin to the monotremes.

"Finally, if the branch of saurians has given rise to the amphibious mammals, it will be most probable that this branch is the source whence all the mammals have taken their origin.

"I therefore believe myself authorized to think that the terrestrial mammals originally descended from those aquatic mammals that we call Amphibia. Because the latter being divided into three branches by the diversity of the habits which, with the lapse of time, they have adopted, some have caused the formation of the Cetacea, others that of the ungulated mammals, and still others that of the unguiculate mammals.

"For example, those of the Amphibia which have preserved the habit of frequenting the shores differ in the manner of taking their food. Some among them accustoming themselves to browse on herbage, such as the morses and lamatines, gradually gave origin to the ungulate mammals, such as the pachyderms, ruminants, etc.; the others, such as the Phocidæ, contracting the habit of feeding on fishes and marine animals, caused the existence of the unguiculate mammals, by means of races which, while becoming differentiated, became entirely terrestrial.

"But those aquatic mammals which would form the habit of never leaving the water, and only rising to breathe at the surface, would probably give origin to the different known cetaceans. Moreover, the ancient and complete habitation of the Cetacea in the ocean has so modified their structure that it is now very difficult to recognize the source whence they have derived their origin.

"Indeed, since the enormous length of time during which these animals have lived in the depths of the sea, never using their hind feet in seizing objects, their disused feet have wholly disappeared, as also their skeleton, and even the pelvis serving as their attachment.

"The alteration which the cetaceans have undergone in their limbs, owing to the influence of the medium in which they live and the habits which they have there contracted, manifests itself also in their fore limbs, which, entirely enveloped by the skin, no longer show externally the fingers in which they end; so that they only offer on each side a fin which contains concealed within it the skeleton of a hand.

"Assuredly, the cetaceans being mammals, it entered into the plan of their structure to have four limbs like the others, and consequently a pelvis to sustain their hind legs. But here, as elsewhere, that which is lacking in them is the result of atrophy brought about, at the end of a long time, by the want of use of the parts which were useless.

"If we consider that in the Phocæ, where the pelvis still exists, this pelvis is impoverished, narrowed, and with no projections on the hips, we see that the lessened (_médiocre_) use of the hind feet of these animals must be the cause, and that if this use should entirely cease, the hind limbs and even the pelvis would in the end disappear.

"The considerations which I have just presented may doubtless appear as simple conjectures, because it is possible to establish them only on direct and positive proofs. But if we pay any attention to the observations which I have stated in this work, and if then we examine carefully the animals which I have mentioned, as also the result of their habits and their surroundings, we shall find that these conjectures will acquire, after this examination, an eminent probability.

"The following _tableau_[191] will facilitate the comprehension of what I have just stated. It will be seen that, in my opinion, the animal scale begins at least by two special branches, and that in the course of its extent some branchlets (_rameaux_) would seem to terminate in certain places.

"This series of animals beginning with two branches where are situated the most imperfect, the first of these branches received their existence only by direct or spontaneous generation.

"A strong reason prevents our knowing the changes successively brought about which have produced the condition in which we observe them; it is because we are never witnesses of these changes. Thus we see the work when done, but never watching them during the process, we are naturally led to believe that things have always been as we see them, and not as they have progressively been brought about.

"Among the changes which nature everywhere incessantly produces in her _ensemble_, and her laws remain always the same, such of these changes as, to bring about, do not need much more time than the duration of human life, are easily understood by the man who observes them; but he cannot perceive those which are accomplished at the end of a considerable time.

"If the duration of human life only extended to the length of a _second_, and if there existed one of our actual clocks mounted and in movement, each individual of our species who should look at the hour-hand of this clock would never see it change its place in the course of his life, although this hand would really not be stationary. The observations of thirty generations would never learn anything very evident as to the displacement of this hand, because its movement, only being that made during half a minute, would be too slight to make an impression; and if observations much more ancient should show that this same hand had really moved, those who should see the statement would not believe it, and would suppose there was some error, each one having always seen the hand on the same point of the dial-plate.

"I leave to my readers all the applications to be made regarding this supposition.

"_Nature_, that immense totality of different beings and bodies, in every part of which exists an eternal circle of movements and changes regulated by law; totality alone unchangeable, so long as it pleases its SUBLIME AUTHOR to make it exist, should be regarded as a whole constituted by its parts, for a purpose which its Author alone knows, and not exclusively for any one of them.

"Each part necessarily is obliged to change, and to cease to be one in order to constitute another, with interests opposed to those of all; and if it has the power of reasoning it finds this whole imperfect. In reality, however, this whole is perfect, and completely fulfils the end for which it was designed."

The last work in which Lamarck discussed the theory of descent was in his introduction to the _Animaux sans Vertèbres_. But here the only changes of importance are his four laws, which we translate, and a somewhat different phylogeny of the animal kingdom.

The four laws differ from the two given in the _Philosophie zoologique_ in his theory (the second law) accounting for the origin of a new organ, the result of a new need.

"_First law_: Life, by its proper forces, continually tends to increase the volume of every body which possesses it, and to increase the size of its parts, up to a limit which it brings about.

"_Second law_: The production of a new organ in an animal body results from the supervention of a new want (_besoin_) which continues to make itself felt, and of a new movement which this want gives rise to and maintains.

"_Third law_: The development of organs and their power of action are constantly in ratio to the employment of these organs.

"_Fourth law_: Everything which has been acquired, impressed upon, or changed in the organization of individuals, during the course of their life is preserved by generation and transmitted to the new individuals which have descended from those which have undergone those changes."

In explaining the second law he says:

"The foundation of this law derives its proof from the third, in which the facts known allow of no doubt; for, if the forces of

## action of an organ, by their increase, further develop this

organ--namely, increase its size and power, as is constantly proved by facts--we may be assured that the forces by which it acts, just originated by a new want felt, would necessarily give birth to the organ adapted to satisfy this new want, if this organ had not before existed.

"In truth, in animals so low as not to be able to _feel_, it cannot be that we should attribute to a felt want the formation of a new organ, this formation being in such a case the product of a mechanical cause, as that of a new movement produced in a part of the fluids of the animal.

"It is not the same in animals with a more complicated structure, and which are able to _feel_. They feel wants, and each want felt, exciting their inner feeling, forthwith sets the fluids in motion and forces them towards the point of the body where an action may satisfy the want experienced. Now, if there exists at this point an organ suitable for this action, it is immediately cited to act; and if the organ does not exist, and only the felt want be for instance pressing and continuous, gradually the organ originates, and is developed on account of the continuity and energy of its employment.

"If I had not been convinced: 1, that the thought alone of an action which strongly interests it suffices to arouse the _inner feeling_ of an individual; 2, that a felt want can itself arouse the feeling in question; 3, that every emotion of _inner feeling_, resulting from a want which is aroused, directs at the same instant a mass of nervous fluid to the points to be set in activity, that it also creates a flow thither of the fluids of the body, and especially nutrient ones; that, finally, it then places in activity the organs already existing, or makes efforts for the formation of those which would not have existed there, and which a continual want would therefore render necessary--I should have had doubts as to the reality of the law which I have just indicated.

"But, although it may be very difficult to verify this law by observation, I have no doubt as to the grounds on which I base it, the necessity of its existence being involved in that of the third law, which is now well established.

"I conceive, for example, that a _gasteropod mollusc_, which, as it crawls along, finds the need of feeling the bodies in front of it, makes efforts to touch those bodies with some of the foremost parts of its head, and sends to these every time supplies of nervous fluids, as well as other fluids--I conceive, I say, that it must result from this reiterated afflux towards the points in question that the nerves which abut at these points will, by slow degrees, be extended. Now, as in the same circumstances other fluids of the animal flow also to the same places, and especially nourishing fluids, it must follow that two or more tentacles will appear and develop insensibly under those circumstances on the points referred to.

"This is doubtless what has happened to all the races of _Gasteropods_, whose wants have compelled them to adopt the habit of feeling bodies with some part of their head.

"But if there occur, among the _Gasteropods_, any races which, by the circumstances which concern their mode of existence or life, do not experience such wants, then their head remains without tentacles; it has even no projection, no traces of tentacles, and this is what has happened in the case of _Bullæa_, _Bulla_, and _Chiton_."

In the _Supplément à la Distribution générale des Animaux_ (Introduction, p. 342), concerning the real order of origin of the invertebrate classes, Lamarck proposes a new genealogical tree. He states that the order of the animal series "is far from simple, that it is branching, and seems even to be composed of several distinct series;" though farther on (p. 456) he adds:

"Je regarde _l'ordre de la production_ des animaux comme formé de deux séries distinctes.

"Ainsi, je soumets à la méditation des zoologistes l'ordre présumé de la _formation_ des animaux, tel que l'exprime le tableau suivant:"

In the matter of the origin of instinct, as in evolution in general, Lamarck appears to have laid the foundation on which Darwin's views, though he throws aside Lamarck's factors, must rest. The "inherited habit" theory is thus stated by Lamarck.

Instinct, he claims, is not common to all animals, since the lowest forms, like plants, are entirely passive under the influences of the surrounding medium; they have no wants, are automata.

"But animals with a nervous system have _wants_, _i.e._, they feel hunger, sexual desires, they desire to avoid pain or to seek pleasure, etc. To satisfy these wants they contract habits, which are gradually transformed into so many propensities which they can neither resist nor change. Hence arise habitual actions and special _propensities_, to which we give the name of _instinct_.

"These propensities are inherited and become innate in the young, so that they act instinctively from the moment of birth. Thus the same habits and instincts are perpetuated from one generation to another, with no _notable_ variations, so long as the species does not suffer change in the circumstances essential to its mode of life."

The same views are repeated in the introduction to the _Animaux sans Vertèbres_ (1815), and again in 1820, in his last work, and do not need to be translated, as they are repetitions of his previously published views in the _Philosophie zoologique_.

Unfortunately, to illustrate his thoughts on instinct Lamarck does not give us any examples, nor did he apparently observe to any great extent the habits of animals. In these days one cannot follow him in drawing a line--as regards the possession of instincts--between the lowest organisms, or Protozoa, and the groups provided with a nervous system.

_Lamarck's meaning of the word "besoins," or wants or needs._--Lamarck's use of the word wants or needs (_besoins_) has, we think, been greatly misunderstood and at times caricatured or pronounced as "absurd." The distinguished French naturalist, Quatrefages, although he was not himself an evolutionist, has protested against the way Lamarck's views have been caricatured. By nearly all authors he is represented as claiming that by simply "willing" or "desiring" the individual bird or other animal radically and with more or less rapidity changed its shape or that of some particular organ or part of the body. This is, as we have seen, by no means what he states. In no instance does he speak of an animal as simply "desiring" to modify an organ in any way. The doctrine of appetency attributed to Lamarck is without foundation. In all the examples given he intimates that owing to changes in environment, leading to isolation in a new area separating a large number of individuals from their accustomed habitat, they are driven by necessity (_besoin_) or new needs to adopt a new or different mode of life--new habits. These efforts, whatever they may be--such as attempts to fly, swim, wade, climb, burrow, etc., continued for a long time "in all the individuals of its species," or the great number forced by competition to migrate and become segregated from the others of the original species--finally, owing to the changed surroundings, affect the mass of individuals thus isolated, and their organs thus exercised in a special direction undergo a slow modification.

Even so careful a writer as Dr. Alfred R. Wallace does not quite fairly, or with exactness, state what Lamarck says, when in his classical essay of 1858 he represents Lamarck as stating that the giraffe acquired its long neck by _desiring_ to reach the foliage of the more lofty shrubs, and constantly stretching its neck for the purpose. On the contrary, he does not use the word "desiring" at all. What Lamarck does say is that--

"The giraffe lives in dry, desert places, without herbage, so that it is obliged to browse on the leaves of trees, and is continually forced to reach up to them. It results from this habit, continued for a long time in all the individuals of its species, that its fore limbs have become so elongated that the giraffe, without raising itself erect on its hind legs, raises its head and reaches six meters high (almost twenty feet)."[192]

We submit that this mode of evolution of the giraffe is quite as reasonable as the very hypothetical one advanced by Mr. Wallace;[193] _i.e._, that a variety occurred with a longer neck than usual, and these "at once secured a fresh range of pasture over the same ground as their shorter-necked companions, and on the first scarcity of food were thereby enabled to outlive them." Mr. Wallace's account also of Lamarck's general theory appears to us to be one-sided, inadequate, and misleading. He states it thus: "The hypothesis of Lamarck--that progressive changes in species have been produced by the attempts of animals to increase the development of their own organs, and thus modify their structure and habits." This is a caricature of what Lamarck really taught. Wants, needs (_besoins_), volitions, desires, are not mentioned by Lamarck in his two fundamental laws (see p. 303), and when the word _besoins_ is introduced it refers as much to the physiological needs as to the emotions of the animal resulting from some new environment which forces it to adopt new habits such as means of locomotion or of acquiring food.

It will be evident to one who has read the original or the foregoing translations of Lamarck's writings that he does not refer so much to mental desires or volitions as to those physiological wants or needs thrust upon the animal by change of circumstances or by competition; and his _besoins_ may include lust, hunger, as well as the necessity of making muscular exertions such as walking, running, leaping, climbing, swimming, or flying.

As we understand Lamarck, when he speaks of the incipient giraffe or long-necked bird as making efforts to reach up or outwards, the efforts may have been as much physiological, reflex, or instinctive as mental. A recent writer, Dr. R. T. Jackson, curiously and yet naturally enough uses the same phraseology as Lamarck when he says that the long siphon of the common clam (Mya) "was brought about by the effort to reach the surface, induced by the habit of deep burial" in its hole.[194]

On the other hand, can we in the higher vertebrates entirely dissociate the emotional and mental activities from their physiological or instinctive acts? Mr. Darwin, in his _Expressions of the Emotions in Man and Animals_, discusses in an interesting and detailed way the effects of the feelings and passions on some of the higher animals.

It is curious, also, that Dr. Erasmus Darwin went at least as far as Lamarck in claiming that the transformations of animals "are in part produced by their own exertions in consequence of their desires and aversions, of their pleasures and their pains, or of irritations or of associations."

Cope, in the final chapter of his _Primary Factors of Organic Evolution_, entitled "The Functions of Consciousness," goes to much farther extremes than the French philosopher has been accused of doing, and unhesitatingly attributes consciousness to all animals. "Whatever be its nature," he says, "the preliminary to any animal movement which is not automatic is an effort." Hence he regards effort as the immediate source of all movement, and considers that the control of muscular movements by consciousness is distinctly observable; in fact, he even goes to the length of affirming that reflex acts are the product of conscious acts, whereas it is plain enough that reflex acts are always the result of some stimulus.

Another case mentioned by Lamarck in his _Animaux sans Vertèbres_, which has been pronounced as absurd and ridiculous, and has aided in throwing his whole theory into disfavor, is his way of accounting for the development of the tentacles of the snail, which is quoted on p. 348.

This account is a very probable and, in fact, the only rational explanation. The initial cause of such structures is the intermittent stimulus of occasional contact with surrounding objects, the irritation thus set up causing a flow of the blood to the exposed parts receiving the stimuli. The general cause is the same as that concerned in the production of horns and other hard defensive projections on the heads of various animals.

In commenting on this case of the snail, Professor Cleland, in his just and discriminating article on Lamarck, says:

"However absurd this may seem, it must be admitted that, unlimited time having been once granted for organs to be developed in series of generations, the objections to their being formed in the way here imagined are only such as equally apply to the theory of their origin by natural selection.... In judging the reasonableness of the second law of Lamarck [referring to new wants, see p. 346] as compared with more modern and now widely received theories, it must be observed that it is only an extension of his third law; and that third law is a fact. The strengthening of the blacksmith's arm by use is proverbially notorious. It is, therefore, only the sufficiency of the Lamarckian hypothesis to explain the first commencement of new organs which is in question, if evolution by the mere operation of forces acting in the organic world be granted; and surely the Darwinian theory is equally helpless to account for the beginning of a new organ, while it demands as imperatively that every stage in the assumed hereditary development of an organ must have been useful.... Lamarck gave great importance to the influence of new wants acting indirectly by stimulating growth and use. Darwin has given like importance to the effects of accidental variations

## acting indirectly by giving advantage in the struggle for existence.

The speculative writings of Darwin have, however, been interwoven with a vast number of beautiful experiments and observations bearing on his speculations, though by no means proving his theory of evolution; while the speculations of Lamarck lie apart from his wonderful descriptive labors, unrelieved by intermixture with other matters capable of attracting the numerous class who, provided they have new facts set before them, are not careful to limit themselves to the conclusions strictly deducible therefrom. But those who read the _Philosophie Zoologique_ will find how many truths often supposed to be far more modern are stated with abundant clearness in its pages." (_Encyc. Brit._, art. "Lamarck.")

COMPARATIVE SUMMARY OF THE VIEWS OF THE FOUNDERS OF THE THEORY OF EVOLUTION, WITH DATES OF PUBLICATION.

-------------+-------------+------------------------+------------+------------ |Erasmus | |Geoffroy St.|Charles Buffon |Darwin |Lamarck |Hilaire |Darwin (1761-1778). |(1790-1794). |(1801-1809-1815). |(1795-1831).|(1859). -------------+-------------+------------------------+------------+------------ | | | | All animals |All animals |All organisms arose from|Unity of |Universal possibly |derived from |germs. First germ |organization|tendency to derived from |a single |originated by |in animal |fortuitous a single |filament. |spontaneous generation. |kingdom. |variability type. | |Development from the | |assumed. | |simple to the complex. |Change of | Time, its | |Animal series not |"milieu | great length,| |continuous, but |ambiant," | stated. | |tree-like; graduated |direct. | | |from monad to man; | | Immutability | |constructed the first | | of species | |phylogenetic tree. | | stated and | | |Founded the |Struggle then denied. |Time, great |Time, great length of, |doctrine of |for |length of, |definitely postulated; |homologies. |existence. Nature |definitely |its duration practically| | advances by |demanded. |unlimited. | | gradations, | | | | passing from | |Uniformitarianism of | | one species | |Hutton and of Lyell |Founder of | to another by| |anticipated. |teratology. | imperceptible| | | | degrees. |Effects of |Effects of favorable |His embryo- | |change of |circumstances, such as |logical | Changes in |climate, |changes of environment, |studies | distribution |direct |climate, soil, food, |influenced | of land and |(briefly |temperature; direct in |his | water as |stated). |case of plants and |philosophic | causing | |lowest animals, indirect|views. | variation. | |in case of the higher | | | |animals and man. | | Effects of | | | | changes of | |Conditions of existence | | climate, | |remaining constant, | | direct. | |species do not vary and | |Competition | |vice-versa. | |strongly Effects of | | | |advocated. changes of | |Struggle for existence; | | food. | |stronger devour the | |Natural |Domesti- |weaker. Competition | |selection. Effects of |cation |stated in case of ai or |Species are | domesti- |briefly |sloth. Balance of |"different |Sexual cation. |referred to. |nature. |modifi- |selection. | | |cations of | Effects of |Effects of |Effects of use and |one and the |Effects of use. (The |use: |disuse, discussed at |same type." |use and only examples|characters |length. | |disuse (in given are the|produced by | | |some callosities |their own |Vestigial structures the| |cases). on legs of |exertions in |remains of organs | | camel, of |consequence |actively used by | | baboon, and |of their |ancestors of present | | the |desires, |forms. | | thickening by|aversions, | | | use of soles |lust, hunger,|New wants or necessities| | on man's |and security.|induced by changes of | | feet.) | |climate, habitat, etc., | | |Sexual |result in production of | | |selection, |new propensities, new | | |law of |habits, and functions. | | |battle. | | | | |Change of habits | | |Protective |originate organs; change| | |mimicry. |of functions create new | | | |organs; formation of new| | |Origin of |habits precede the | | |organs before|origin of new or | | |development |modification of organs | | |of their |already formed. | | |functions. | | | | |Geographical isolation | |Isolation |Inheritance |suggested as a factor in| |"an |of acquired |case of man. | |important |characters | | |element." |(vaguely |Swamping effects of | | |stated). |crossing. | | | | | | |Instincts |Lamarck's definition of | | |result of |species the most | | |imitation. |satisfactory yet stated.| | | | | | |Opposed |Inheritance of acquired | |Inheritance |preformation |characters. | |of acquired |views of | | |characters. |Haller and |Instinct the result of | | |Bonnet. |inherited habits. | | | | | | | |Opposed preformation | | | |views; epigenesis | | | |definitely stated and | | | |adopted. | | | | | | -------------+-------------+------------------------+------------+------------

FOOTNOTES:

[179] [Cabanis.] _Rapp. du Phys. et du Moral de l'Homme_, pp. 38 à 39, et 85.

[180] Lamarck's idea of the animal series was that of a branched one, as shown by his genealogical tree on p. 193, and he explains that the series begins at least by two special branches, these ending in branchlets. He thus breaks entirely away from the old idea of a continuous ascending series of his predecessors Bonnet and others. Professor R. Hertwig therefore makes a decided mistake and does Lamarck a great injustice in his "Zoölogy," where he states: "Lamarck, in agreement with the then prevailing conceptions, regarded the animal kingdom as a series grading from the lowest primitive animal up to man" (p. 26); and again, on the next page, he speaks of "the theory of Geoffroy St.-Hilaire and Lamarck" as having in it "as a fundamental error the doctrine of the serial arrangement of the animal world" (English Trans.). Hertwig is in error, and could never have carefully read what Lamarck did say, or have known that he was the first to throw aside the serial arrangement, and to sketch out a genealogical tree.

[181] The foregoing pages (283-286) are reprinted by the author from the _Discours_ of 1803. See pp. 266-270.

[182] Perrier thus comments on this passage: "_Ici nous sommes bien près, semble-t-il, non seulement de la lutte pour la vie telle one la concevra Darwin, mais même de la sélection naturelle. Malheureusement, au lieu de poursuivre l'idée, Lamarck aussitôt s'engage dans une autre voie_," etc. (_La Philosophie zoologique avant Darwin_, p. 81).

[183] The expression "_sentiment intérieur_" may be nearly equivalent to the "organic sense" of modern psychologists, but more probably corresponds to our word consciousness.

[184] Lamarck's division of _Animaux sensibles_ comprises the insects, arachnids, crustacea, annelids, cirrhipedes, and molluscs.

[185] Rather a strange view to take, as the brain of insects is now known to be nearly as complex as that of mammals.

[186] Richerand, _Physiologie_. vol ii. p. 151.

[187] "As all animals do not have the power of performing voluntary acts, so in like manner _instinct_ is not common to all animals: for those lacking the nervous system also want the organic sense, and can perform no instinctive acts.

"These imperfect animals are entirely passive, they do nothing of themselves, they have no wants, and nature as regards them treats them as she does plants. But as they are irritable in their parts, the means which nature employs to maintain their existence enables them to execute movements which we call actions."

It thus appears that Lamarck practically regards the lowest animals as automata, but we must remember that the line he draws between animals with and without a nervous system is an artificial one, as some of the forms which he supposed to be destitute of a nervous system are now known to possess one.

[188] It should be noticed that Lamarck does not absolutely state that there are no variations whatever in instinct. His words are much less positive: "_Sans offrer de variation notable._" This dues not exclude the fact, discovered since his time, that instincts are more or less variable, thus affording grounds for Darwin's theory of the origin of new kinds of instincts from the "accidental variation of instincts." Professor James' otherwise excellent version of Lamarck's view is inexact and misleading when he makes Lamarck say that instincts are "perpetuated _without variation_ from one generation to another, so long as the outward conditions of existence remain the same" (_The Principles of Psychology_, vol. ii., p. 678, 1890). He leaves out the word notable. The italics are ours. Farther on (p. 337), it will be seen that Lamarck acknowledges that in birds and mammals instinct is variable.

[189] It is interesting to compare with this Darwin's theory of the origin of the same animals, the flying squirrels and Galeopithecus (_Origin of Species_, 5th edition, New York, pp. 173-174), and see how he invokes the Lamarckian factors of change of "climate and vegetation" and "changing conditions of life," to originate the variations before natural selection can act. His account is a mixture of Lamarckism with the added Darwinian factors of competition and natural selection. We agree with this view, that the change in environment and competition sets the ball in motion, the work being finished by the selective process. The act of springing and the first attempts at flying also involve strong emotions and mental efforts, and it can hardly be denied that these Lamarckian factors came into continual play during the process of evolution of these flying creatures.

[190] This sagacious, though crude suggestion of the origin of birds and mammals from the reptiles is now, after the lapse of nearly a century, being confirmed by modern morphologists and palæontologists.

[191] Reproduced on page 193.

[192] This is taken from my article, "Lamarck and Neo-lamarckianism," in the _Open Court_, Chicago, February, 1897. Compare also "Darwin Wrong," etc., by R. F. Licorish, M.D., Barbadoes, 1898, reprinted in _Natural Science_, April, 1899.

[193] _Natural Selection_, pp. 41-42.

[194] _American Naturalist_, 1891, p. 17.

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