Chapter XVI

. and the Pedigree, p. 398.)

At the time when Goethe in this way sketched the fundamental features of the Theory of Descent, another German philosopher, Gottfried Reinhold Treviranus, of Bremen (born 1776, died 1837), was zealously engaged at the same work. As Wilhelm Focke has recently shown, Treviranus, even in the earliest of his greater works, “The Biology or Philosophy of Animate Nature,” which appeared at the beginning of the present century, had already developed monistic views of the unity of nature, and of the genealogical connection of the species of organisms, which entirely correspond with our present view of the matter. In the first three volumes of the Biology, which appeared successively in 1802, 1803, and 1805 (therefore several years before Oken’s and Lamarck’s principal works), we find numerous passages which are of interest in this respect. I shall here quote only a few of the most important.

In speaking of the principal question of our theory, the question of the origin of organic species, Treviranus makes the following remarks:—“Every form of life can be produced by physical forces in one of two ways: either by coming into being out of formless matter, or by modification of an already existing form by a continued process of shaping. In the latter case the cause of this modification may lie either in the influence of a dissimilar male generative matter upon the female germ, or in the influence of other powers which operate only after procreation. In every living being there exists the capability of an endless variety of form-assumption; each possesses the power to adapt its organization to the changes of the outer world, and it is this power put into action by the change of the universe that has raised the simple zoophytes of the primitive world to continually higher stages of organization, and has introduced a countless variety of species into animate nature.”

By _zoophytes_, Treviranus here means organisms of the lowest order and of the simplest character, namely, those neutral primitive beings which stand midway between animals and plants, and on the whole correspond with our _protista_. “These zoophytes,” he remarks in another passage, “are the original forms out of which all the organisms of the higher classes have arisen by gradual development. We are further of opinion that every species, as well as every individual, has certain periods of growth, of bloom, and of decay, but that the decay of a species is _degeneration_, not dissolution, as in the case of the individual. From this it appears to us to follow that it was not the great catastrophes of the earth (as is generally supposed) which destroyed the animals of the primitive world, but that many survived them, and it is more probable that they have disappeared from existing nature, because the species to which they belonged have completed the circle of their existence, and have become changed into other kinds.”

When Treviranus, in this and other passages, points to _degeneration_ as the most important cause of the transformation of the animal and vegetable species, he does not understand by it what is now commonly called degeneration. With him “degeneration” is exactly what we now call _Adaptation_ or _modification_, by the action of external formative forces. That Treviranus explained this trans-transformation of organic species by Adaptation, and its preservation by Inheritance, and thus the whole variety of organic forms by the interaction of Adaptation and Inheritance, is clear also from several other passages. How profoundly he grasped the mutual dependence of all living creatures on one another, and in general the _universal connection between cause and effect_—that is, the monistic causal connection between all members and parts of the universe—is further shown, among others, by the following remarks in his Biology:—“The living individual is dependent upon the species, the species upon the fauna, the fauna upon the whole of animate nature, and the latter upon the organism of the earth. The individual possesses indeed a peculiar life, and so far forms its own world. But just because its life is limited it constitutes at the same time an organ in the general organism. Every living body exists in consequence of the universe, but the universe, on the other hand, exists in consequence of it.”

It is self-evident that so profound and clear a thinker as Treviranus, in accordance with this grand mechanical conception of the universe, could not admit for man a privileged and exceptional position in nature, but assumed his gradual development from lower animal forms. And it is equally self-evident, on the other hand, that he did not admit a chasm between organic and inorganic nature, but maintained the absolute unity of the organization of the whole universe. This is specially attested by the following sentence:—“Every inquiry into the influence of the whole of nature on the living world must start from the principle, that all living forms are products of physical influences, which are acting even now, and are changed only in degree, or in their direction.” Hereby, as Treviranus himself says, “The fundamental problem of biology is solved,” and we add, solved in a purely mechanical or monistic sense.

Neither Treviranus nor Goethe is commonly considered the most eminent of the German nature-philosophers, but Lorenz Oken, who, in establishing the vertebral theory of the skull, came forward as a rival to Goethe, and did not entertain a very kindly feeling towards him. Although they lived for some time in the same neighbourhood, yet the natures of these two men were so very different, that they could not well be drawn towards each other. Oken’s “Manual of the Philosophy of Nature,” which may be designated as the most important production of the nature-philosophy school then existing in Germany, appeared in 1809, the same year in which Lamarck’s fundamental work, the “Philosophie Zoologique,” was published. As early as 1802, Oken had published an “Outline of the Philosophy of Nature.” As we have already intimated, in Oken’s as in Goethe’s works, a number of valuable and profound thoughts are hidden among a mass of erroneous, very eccentric, and fantastic conceptions. Some of these ideas have only quite recently and gradually become recognized in science, many years after they were first expressed. I shall here quote only two thoughts, which are almost prophetic, and which at the same time stand in the closest relation to the theory of development.

One of the most important of Oken’s theories, which was formerly very much decried, and was most strongly combatted, especially by the so-called “exact experimentalists,” is the idea that the phenomena of life in all organisms proceed from a common chemical substance, so to say, from a general simple _vital-substance_, which he designated by the name _Urschleim_, or _original slime_. By it he meant, as the name indicates, a mucilaginous substance, an albuminous combination, which exists in a semi-fluid condition of aggregation, and possesses the power, by adaptation to different conditions of existence in the outer world and by interaction with its material, of producing the most various forms. Now, we need only change the expression “original slime” (Urschleim) into _Protoplasm_, or _cell-substance_, in order to arrive at one of the grandest results which we owe to microscopic investigations during the last ten years, more especially to those of Max Schultze. By these investigations it has been shown that in all living bodies, without exception, there exists a certain quantity of mucilaginous albuminous matter, in a semi-fluid condition; and that this nitrogen-holding carbon-compound is exclusively the original seat and agent of all the phenomena of life, and of all production of organic forms. All other substances which appear in the organism, besides these, are either formed by this active matter of life, or have been introduced from without. The organic egg, the original cell out of which every animal and plant is first developed, consists essentially only of one round little lump of such albuminous matter. Even the yolk of an egg is nothing but albumen, mixed with granules of fat. Oken was therefore right when, more divining than knowing, he made the assertion—“Every organic thing has arisen out of slime, and is nothing but slime in different forms. This primitive slime originated in the sea, from inorganic matter in the course of planetary-evolution.”

Another equally grand idea of the same philosopher is closely connected with his theory of primitive slime, which coincides with the extremely important _Protoplasm theory_. For Oken, as early as 1809, asserted that the primitive slime produced in the sea by spontaneous generation, at once assumed the form of microscopically small bladders, which he called “_Mile_,” or “_Infusoria_.” “Organic nature has for its basis an infinity of such vesicles.” These little bladders arise from original semi-fluid globules of the primitive slime, by the fact of their periphery becoming condensed. The simplest organism, as well as every animal and every plant of higher kind, is nothing else than “an accumulation (synthesis) of such infusorial bladders, which by various combinations assume various forms, and thus develop into higher organisms.” Here again we need only translate the expression _little bladder_, or _infusorium_, by the word _cell_, and we arrive at the Cell theory, one of the grandest biological theories of our century. Schleiden and Schwann, about thirty years ago, were the first to furnish experiential proof that all organisms are either simple cells, or accumulations (syntheses) of such cells, and the more recent protoplasm theory has shown that protoplasm (the original slime) is the most essential (and sometimes the only) constituent part of the genuine cell. The properties which Oken ascribes to his Infusoria are exactly the properties of cells, the properties of elementary beings, by whose accumulation, combination, and varying development, the higher organisms are formed.

These two extremely fruitful thoughts of Oken, on account of the absurd form in which he expressed them, were at first little heeded, or entirely misunderstood, and it was reserved for a much later era to establish them by actual observation. The supposition that the individual species of plants and animals originated from common prototypes by a slow and gradual development of the higher organisms out of lower ones, was of course most closely connected with these ideas. Man’s descent from lower organisms was likewise asserted by Oken—“Man has been developed, not created.” Although many arbitrary perversities and extravagant fancies may be found in Oken’s philosophy of nature, they must not prevent us paying our just admiration to these grand ideas, which were so far in advance of their age. This much is clearly evident from the statements of Goethe and Oken which we have quoted, and from the views of Lamarck and Geoffroy which have to be discussed next, that during the first decade of our century no doctrine approached so nearly to the natural Theory of Descent, newly established by Darwin, as the much decried “Natur-philosophie.”

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